The top molar cusps are seemingly bunoselenomorph

The top molar cusps are seemingly bunoselenomorph

He has got an initial and you may straight postprotocrista labially mainly based. The preprotocrista, relatively upright, satisfies a well-create parastyle. New metaconule try somewhat quicker that’s large on M3. The fresh new premetaconulecrista while the fossae are simple as opposed to neocrista. The newest rear fossa try tilted, particularly towards the M3. From upright into the M1, the new postmetaconulecrista will get sigmoidal to the M3, skipping the newest metacone. The newest labial cusps have the same axis which have relatively upright cristae. New paracone labial rib try outstanding, but without anterior groove. There’s no metacone labial rib. The latest para-, meso-, and you can metastyles try salient. The lingual cingulum is quite weakened so you’re able to absent such as the entostyle.

The newest recorded build of your next lower premolar is common to possess derived stalk pecoran and you will crown pecoran into the which have a well-build mesolingual conid and you can rear cristid , . Additionally, the current presence of an enamel retailer to your p1 excludes an enthusiastic attribution in order to Prodremotherium or Mosaicomeryx . Even if the white teeth aren’t while the bunoselenodont as with Amphitragulus elegans, the fresh new known specimens have very primitive has, such as for example a highly faster metastylid, a decreased top, a small postentocristid, and even though laterally compacted, bulged lingual cuspids on the lower molars. Each one of these services prohibit an association so you can either Dremotherium or Bedenomeryx , . Additionally, the new most small-size away from “Amphitragulus” quercyi was a distinguishing element with the latest Oligocene ruminants out of European countries , , , .

Dremotherium cf. guthi out-of Gaimersheim1: step one kept maxillary bones towards the enamel line M1-M3 (SNSB-BSPG 1952II4855). out of Mailhat: dos left P4-M2 (MNHN LIM154). Babameryx engesseri off Gaimersheim1: step three left fragmentary mandible which have m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you may (c) occlusal viewpoints. The dimensions bar try dos cm.

Holotype known specimens guidance

Holotype. Jehenne (1987) has actually appointed the newest specimen IPHEP LM.1969.MA.forty given that holotype. This can be a partial right maxillary bones with P3-M3 (, Fig. 2), away from La Milloque (MP29, France).

Referred specimens. Mailhat MNHN LIM154 kept P4-M2 (it functions, Fig. six.2); Gaimersheim1 SNSB-BSPG 1952II4855 leftover maxillary having M1-M3 (that it functions, Fig. 6.1); SNSB-BSPG 1952II4870 correct m2.


All the way down permanent dentition. The reduced molar is extremely badly preserved. However, so it huge ruminant possesses a completely selenomorph trigonid. The cristid try elongated and also the cuspids is large to the enamel slightly pleated. The metastylid are well developed.

Top long lasting dentition. Brand new P4 are defectively kept. The lingual region try shed. The anterolabial cone features a failure labial rib. The fresh new prior stylid are damaged. The fresh new rear a person is quick but introduce. No neocrista obstructs the latest fossa. The fresh quadratic upper molars when you look at the occlusal examine incur a properly-created metaconule some smaller compared to brand new protocone. The fresh new cusps are particularly selenomorph. The preprotocrista in addition to postmetaconulecrista is labially centered and some straight. This new postprotocrista is long, highly rounded, and you can labially created. It could be completely separated (SNSB-BSPG 1952II4855) or fused (MNHN LIM154) with the premetaconulecrista. Aforementioned finishes at axis of your mesostyle. The fresh rear basin can have a little metaconule bend toward M2, huge on the M3. Brand new paracone, having a good paracone rib, in addition to metacone, labially flat and you may concave, is actually aimed. The newest mesostyle and you will parastyle are create. They setting round quick articles. The brand new mesostyle try large out-of M1 so you can M3. The latest entostyle is actually weakened or absent. There is none a great lingual cingulum related new protocone nor a great metastyle forming a highly-arranged cuspid. But not, brief prior and you can posterior cingula are present.


This ruminant is fairly complex in becoming better-selenodont and achieving zero lingual cingula in the upper molars, usually contained in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and Babameryx , , , [61–63], . In contrast to Bachitherium, this variety lacks an enthusiastic anterior groove towards the very large labial paracone rib , , . The fresh studied fossils be much more similar to the state-of-the-art base Pecora and this new ancient top Pecora (such as Amphitragulus, Dremotherium, Bedenomeryx). The enamel are slightly pleated, whereas it’s essentially simple into the Prodremotherium. Also, how big is these teeth is significantly larger than inside Prodremotherium on the dated Quercy collections and you can Pech Desse . Tooth become more selenomorph than in “Amphitragulus” and you may Bedenomeryx , , additionally the parastyle isn’t that anteriorly projected than observed for the “Amphitragulus” feningrei , , . Looking at the neodiagnosis suggested because of the Mennecart , these fossils is all the showing popular features of Dremotherium. Brand new currently laid out variety within the European countries try D. feignouxi (sorts of variety) and you may D. guthi , , . The latter try some smaller compared to the sort kinds , just like the fossils revealed here. D. feignouxi just took place within the Agenian (MN1-MN2; ), whereas D. guthi resided for the newest Oligocene (MP28-MP30; ). Yet not, from inside the absence of extra better-maintained point, a definitive assignation for the varieties is not possible.


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